1981 | School of Medicine, Kumamoto University, M.D. |
1986 | Graduate School, Division of Medicine, Kyoto University, Ph.D. |
1986 | Post-doctoral fellow, University of California, Los Angeles |
1989 | Assistant Professor, School of Medicine, Yamaguchi University |
1990 | Assistant Professor, School of Medicine, Tokyo Medical and Dental University |
1992 | Lecturer, School of Medicine, Tokyo Medical and Dental University |
1994 | Associate Professor, School of Medicine, Tokyo Medical and Dental University |
1999 | Professor, School of Medicine, Tohoku University |
2004 | Professor, Institute for Virus Research, Kyoto University |
2014 | Director, Institute for Virus Research, Kyoto University |
2016 | Deputy Director, Institute for Frontier Life and Medical Sciences, Kyoto University |
2018 | Director, Institute for Frontier Life and Medical Sciences, Kyoto University |
2022 | Program-Specific Professor, Institute for Liberal Arts and Sciences, Kyoto University |
Immunology, Molecular biology, Virology
The Japanese Society for Virology, The Japanese Society for AIDS Research